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70岁的寡妇托娃·沙利文在索维尔湾水族馆做夜班清洁工，三十年前儿子的失踪一直是她心中的痛。她与馆里聪明的太平洋巨型章鱼马塞勒斯建立起奇妙的羁绊，马塞勒斯总能在她难过时给予无声的陪伴。与此同时，迷茫的青年卡梅隆·卡斯莫正在寻找自己的生父，他的人生也因这场寻找而逐渐清晰。两条看似无关的故事线，因马塞勒斯的意外发现慢慢交织。

　　Shipwreck survivors are found on Beiru Island (Infanto tô), which was previously used for atomic tests. The interior is amazingly free of radiation effects, and they believe that they were protected by a special juice that was given to them by the island's residents. A joint expedition of Rolisican and Japanese scientists explores Beiru and discovers many curious things, including two women only one foot (30 centimeters) high. Unscrupulous expedition leader Clark Nelson abducts the women and puts them in a vaudeville show. But their sweet singing contains a telepathic cry for help to Mothra, a gigantic moth that is worshiped as a deity by the island people. The giant monster heeds the call of the women and heads to Tokyo, wreaking destruction in its path.

濒临破产的老板李大顺（李孟武 饰）为逆转绝境，与死磕项目的技术宅李谱（肖轶 饰）踏上寻“小分子水”之路，临时搭挡的组合囧况百出，还没发车就透露着热闹，并偶遇了身世可怜的少年六六……